PythonとFlaskをLolipopで動かす際のエラー

Python,Flaskを用いたプログラム

上記の通り、Lolipopでブログのようなプログラムを動かしたいのですが、
Internal Server Errorが出てしまいます。
最小限のコードでは動きましたが、オリジナルの物をアップロードするとエラーになってしまいます
解決策を提示していただけると幸いです。

エラーメッセージ

Internal Server Error
The server encountered an internal error or misconfiguration and was unable to complete your request.
Please contact the server administrator at https://lolipop.jp/support/ to inform them of the time this error occurred, and the actions you performed just before this error.
More information about this error may be available in the server error log.
Additionally, a 500 Internal Server Error error was encountered while trying to use an ErrorDocument to handle the request.

該当のソースコード（すべて貼っておきます）

python
1#views.py
2from typing import Text
3from flask import Flask,request,redirect,url_for,render_template,flash,session
4from sqlalchemy.sql.functions import user
5from models.entries import Entry ,SESSION,User
6from sqlalchemy.engine import base
7from sqlalchemy import text
8from datetime import datetime
9
10app = Flask(__name__)
11app.config['DEBUG'] = True
12app.config['USERNAME'] = 'admin'
13app.config['PASSWORD'] = 'admin'
14app.config['SECRET_KEY'] = 'gmowyu7938q2y7uhfsiu7827rngw7390167reidnsdhgoi2yifg6t783hj4i'
15
16@app.route('/')
17def show_entries():
18    if not session.get('logged_in'):
19        return redirect(url_for('login')) 
20    entrie = SESSION.query(Entry).all()
21    usernames = session.get('username')
22    return render_template('home.html',entries = entrie,usernames = usernames)
23@app.route('/signup', methods=['GET', 'POST'])
24def signup(): 
25    if request.method == 'POST':
26        userm = User(
27            name = request.form['usernames'],
28            password = request.form['passwords'],
29        )
30        SESSION.add(userm)
31        SESSION.commit()
32        flash('signuped')
33        return redirect(url_for('login'))
34    elif request.method == 'GET':
35        return render_template('signups.html')
36    else:
37        pass
38@app.route('/login',methods=['GET','POST'])
39def login():
40    if request.method == 'POST':
41        pname = request.form['username']
42        p1 = SESSION.query(User).filter(text("name=:name")).params(name=pname)#.first()
43        
44        for row in p1:
45            NAME = row.name
46            PASSWD = row.password
47        if request.form['username'] != NAME:
48            flash('error:username')
49        elif request.form['password'] != PASSWD:
50            flash('error:passwd')
51        else:
52            session['logged_in']=True
53            session['username']=NAME
54            flash('logged_in')
55            return redirect(url_for('show_entries'))
56    return render_template('login.html')
57@app.route('/logout')
58def logout():
59    session.pop('logged_in',None)
60    session.pop('username',None)
61    flash('logged out')
62    return redirect(url_for('show_entries')) 
63@app.route('/entries',methods=['POST'])
64def add_entry():
65    if not session.get('logged_in'):
66        return redirect(url_for('login')) 
67    entry = Entry(
68        long = request.form['long'],
69        subject = request.form['subject'],
70        comment = request.form['comment'],
71        created_at = datetime.utcnow()
72    )
73    SESSION.add(entry)
74    SESSION.commit()
75    flash('created')
76    return redirect(url_for('show_entries')) 
77@app.route('/entries/new')
78def new_entry():
79    if not session.get('logged_in'):
80        return redirect(url_for('login')) 
81    return render_template('new.html')

python
1#entries.py
2from sqlalchemy.sql.sqltypes import DateTime, Time
3from sqlalchemy import create_engine
4from sqlalchemy.ext.declarative import declarative_base
5from sqlalchemy import Column, Integer, String
6from sqlalchemy.orm import sessionmaker
7
8engine = create_engine('sqlite:///coco.db', echo=True , connect_args={'check_same_thread': False}) #, timeout=10, uri=True , check_same_thread=True
9Base = declarative_base()
10class Entry(Base):
11    __tablename__ = 'entries'
12    id = Column(Integer,primary_key=True)
13    long = Column(String(10))
14    subject = Column(String(50))
15    comment = Column(String(50))
16    created_at = Column(DateTime)
17    class User(Base):
18    __tablename__ = 'user'
19    id = Column(Integer, primary_key=True)
20    name = Column(String)
21    password = Column(String)
22    roll = Column(String)
23    Session = sessionmaker(bind=engine)
24SESSION = Session()
25Base.metadata.create_all(engine)
26userdata = User(
27    name = "admin",
28    password = "admin",
29    roll = True
30)
31SESSION.add(userdata)
32SESSION.commit()

python
1#config.py
2DEBUG=True
3USERNAME='admin'
4PASSWORD='admin'
5SECRET_KEY = 'f47wyhdfa8387hadfja3798293uhda83jk'
6SQLALCHEMY_DATABASE_URI = 'sqlite:///coco.db'
7SQLALCHEMY_TRACK_MODIFICATIONS = True

python
1#!/usr/local/bin/python3.7
2#index.cgi
3from wsgiref.handlers import CGIHandler
4from views import app
5CGIHandler().run(app)

#.htaccess
RewriteEngine On
RewriteCond %{REQUEST_FILENAME} !-f
RewriteRule ^(.*)$ /ffftp/index.cgi/$1 [QSA,L]

html
1<!--home.html-->
2{% extends "layout.html" %}
3{% block content %}
4<ul class="list-group list-group-flush">
5    {% for entry in entries %}
6    <div class="card">
7        <div class="card-body">
8            <h5 class="card-title">{{ entry.id }}:{{ usernames }}さんが、{{ entry.subject }}を{{ entry.long }}分学習しました！</h5><!--{{ usernames }}さんが、-->
9            <h6>一言:{{ entry.comment }}</h6>
10            <h7>created-at:{{ entry.created_at }}</h7>
11        </div>
12    </div>
13    {% else %}
14    noarticle
15    {% endfor %}
16</ul>
17{% endblock %}

html
1<!--layout.html-->
2<!DOCTYPE html>
3<title>coco study</title>
4<link rel="stylesheet" href="https://maxcdn.bootstrapcdn.com/bootstrap/4.0.0/css/bootstrap.min.css">
5<script src="https://code.jquery.com/jquery-3.2.1.slim.min.js"></script>
6<script src="https://cdnjs.cloudflare.com/ajax/libs/popper.js/1.12.9/umd/popper.min.js"></script>
7<script src="https://maxcdn.bootstrapcdn.com/bootstrap/4.0.0/js/bootstrap.min.js"></script>
8<div class="container">
9    <nav class="navbar navbar-expand-lg navbar-light bg-light">
10        <a class="navbar-brand" href="{{url_for('show_entries')}}">Coco study</a>
11        <button class="navbar-toggler" type="button" data-toggle="collapse" data-target="#navbarNav" aria-controls="navbarNav" aria-expanded="false" aria-label="Toggle navigation">
12            <span class="navbar-toggler-icon"></span>
13        </button>
14        <div class="collapse navbar-collapse" id="navbarNav">
15            <ul class="nav navbar-nav navbar-right">
16                {% if not session.logged_in %}
17                <li class="nav-item">
18                    <a class="nav-link" href="{{url_for('login')}}">login</a>
19                </li>
20                <li class="nav-item">
21                    <a class="nav-link" href="{{url_for('signup')}}">signup</a>
22                </li>
23                {% else %}
24                <li class="nav-item">
25                    <a class="nav-link" href="{{url_for('new_entry')}}">new entry</a>
26                </li>
27                <li class="nav-item">
28                    <a class="nav-link" href="{{url_for('logout')}}">logout</a>
29                </li>
30                {% endif %}
31            </ul>
32        </div>
33    </nav>
34    {% for message in get_flashed_messages() %}
35        <div class="alert alert-info" role="alert">
36            {{ message }}
37        </div>
38    {% endfor %}
39    <div class="blog-body">
40        {% block content %}
41        {% endblock %}
42    </div>
43</div>

html
1<!--login.html-->
2{% extends "layout.html" %}
3{% block content %}
4<form action="/login" method="POST">
5    <div class="form-group">
6        <label for="InputTitle">username</label>
7        <input type="text" class="form-control" id="InputTitle" name="username">
8    </div>
9    <div class="form-group">
10        <label for="InputPassword">password</label>
11        <input type="password" class="form-control" id="InputPassword" name="password">
12    </div>
13    <button type="submit" class="btn btn-primary">login</button>
14</form>
15{% endblock %}

html
1<!--new.html-->
2{% extends "layout.html" %}
3{% block content %}
4<form action="{{url_for('add_entry')}}" method="POST" class="add-entry">
5    <div class="form-group">
6        <label for="InputSubject">Subject</label>
7        <input type="text" class="form-control" id="InputSubject" name="subject">
8    </div>
9    <div class="form-group">
10        <label for="Inputlong">long</label>
11        <input type="text" class="form-control" id="Inputlong" name="long">
12        <label for="Inputlong">分</label>
13    </div>
14    <div class="form-group">
15        <label for="inputcomment">comment</label>
16        <input type="text" class="form-control" id="Inputcomment" name="comment">
17    </div>
18    <button type="submit" class="btn btn-primary">create</button>
19</form>
20{% endblock %}

html
1<!--signups.html-->
2{% extends "layout.html" %}
3{% block content %}
4<form action="/signup" method="POST">
5    <div class="form-group">
6        <label for="Inputname">username</label>
7        <input type="text" class="form-control" id="Inputname" name="usernames">
8    </div>
9    <div class="form-group">
10        <label for="InputPassword">password</label>
11        <input type="password" class="form-control" id="InputPassword" name="passwords">
12    </div>
13    <button type="submit" class="btn btn-primary">signup</button>
14</form>
15{% endblock %}

試したこと

lolipopのパス、パーミッション、htaccess、index.cgiの修正

補足情報

ディレクトリ構造

/|
 |-/models
        |-__init__.py(空)
        |-entries.py
 |-/templates
        |-home.html
        |-layout.html
        |-login.html
        |-new.html
        |-signups.html
 |-.htaccess
 |-__init__.py(空)
 |-coco.db
 |-config.py
 |-index.cgi
 |-views.py